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See citation statistics at my Google Scholar page here.

Xu, C., Silliman, B. R., Chen, J., Li, X., Thomsen, M. S., Zhang, Q., Lee, J., Lefcheck, J. S., Daleo, P., Hughes, B. B., Jones, H. P., Wang, R., Wang, S., Smith, C., Xi, X., Altieri, A. H., van de Koppel, J., Palmer, T. M., Liu, L., Wu, J., Li, B., He, Q.* (2023) Herbivory limits success of vegetation restoration globally. Science, 382, 589-594.

          Featured on the cover

          Subject of a Perspective article at Science

He, Q.* (2023). A drowned future for coastal ecosystems. Nature, 621, 44-45.

Li, X., Wang, H., McCauley, D. J., Altieri, A. H., Silliman, B. R., Lefcheck, J. S., Wu, J., Li, B., He, Q.* (2023). A wide megafauna gap undermines China’s expanding coastal ecosystem conservation. Science Advances, 9(32), eadg3800.

          Subject of a News article at Nature

Zhang, K., Yan, J., He, Q., Xu, C., van de Koppel, J., Wang, B., ... & Liu, Q. X. (2023). Self-organized mud cracking amplifies the resilience of an iconic “Red Beach” salt marsh. Science Advances, 9(18), eabq3520.

Chen, J., Li, C., Wu, C., Sun, X., Feng, X., Zhao, J., Ma, Z., Wu, J., Bertness, M.D., Li, B., He, Q.* (2023). Top-down control of macrofauna: Are waterbirds passengers or drivers in wetlands?. Biological Conservation, 279, 109903.

Thomsen M S, A H Altieri, C Angelini, M J Bishop, F Bulleri, R Farhan, V M M Frühling, P E Gribben, S B Harrison, Q He*, M Klinghardt, J Langeneck, B S Lanham, L Mondardini, Y Mulders, S Oleksyn, A P Ramus, D R Schiel, T Schneider, A Siciliano, B R Silliman, D A Smale, P M South, T Wernberg, S Zhang, G Zotz. 2022. Heterogeneity within and among co-occurring foundation species increases biodiversity. Nature Communications, 13, 581

Wang, B., Zhang, K., Liu, Q. X., He, Q., van de Koppel, J., Teng, S. N., ... & Xu, C. (2022). Long-distance facilitation of coastal ecosystem structure and resilience. Proceedings of the National Academy of Sciences USA, 119(28), e2123274119.

Ma, S., Ren, J., Wu, C., Cheng, F., Wang, X., Li, B., & He, Q.* (2022). Hydrological control of threshold transitions in vegetation over early-period wetland development. Journal of Hydrology, 610, 127931.

Ren J, Chen J, Xu C, van de Koppel J, Thomsen MS, Qiu S, Cheng F, Song W, Liu QX, Xu C, Bai J, Zhang Y, Cui B, Bertness MD, Silliman BR, Li B, He Q*. 2021. An invasive species erodes the performance of coastal wetland protected areas. Science Advances, 7: eabi8943.

Li C, H Wang, X Liao, R Xiao, K Liu, J Bai, B Li, Q He*. 2022. Heavy metal pollution in coastal wetlands: a systematic review of studies globally over the past three decades. Journal of Hazardous Materials, 424, 127312

Pennings SC, & He, Q. 2021. Community ecology of salt marshes. In: D FitzGerald, Z Hughes (eds). Salt Marshes: Function, Dynamics, and Stresses. Cambridge University Press, Cambridge, UK.

Qian W, J Chen, Q Zhang, C Wu, Q Ma, BR Silliman, J Wu, B Li, Q He*. 2021. Top-down control of foundation species recovery during coastal wetland restoration. Science of the Total Environment, 769, 144854.

Wang S, Q He, Y Zhang, Q Sheng, B Li, J Wu. 2021. Habitat‐dependent impacts of exotic plant invasions on benthic food webs in a coastal wetland. Limnology and Oceanography, 66, 1256-1267.

Miao X, Zhang H, Xia W, He Q, Wen T, Wang B, Teng SN, Liu Q-X, Niu K, Xu C. 2021. Can we use plant performance as indicators to infer plant-crab interactions in coastal saltmarshes? Ecological Indicators, 129, 107911.

He Q*. 2020. Beauty over the muck: appreciating wetlands via the tales of people. Current Biology 30 (3), R101-R102.

He Q*, H Li, C Xu, Q Sun, MD Bertness, C Fang, B Li, BR Silliman. 2020. Consumer regulation of the carbon cycle in coastal wetland ecosystems. Philosophical Transactions of the Royal Society B 375 (1814), 20190451. 

Qiu S, S Liu, S Wei, X Cui, M Nie, J Huang, Q He, RT Ju, B Li. 2020. Changes in multiple environmental factors additively enhance the dominance of an exotic plant with a novel trade‐off pattern. Journal of Ecology 108 (5), 1989-1999.

He, Q., & Silliman, B. R. 2019. Climate change, human impacts, and coastal ecosystems in the Anthropocene. Current Biology 29: R1021-R1035.

He, Q. 2019. Conservation: ‘No net loss’ of wetland quantity and quality. Current Biology 29: R1070-R1072.

He, Q., Silliman, B. R., van de Koppel, J. & Cui, B. 2019. Weather fluctuations affect the impact of consumers on vegetation recovery following a catastrophic die-off. Ecology 100: e02559.

Silliman, B.R., He, Q., Angelini, C., Smith, C.S., Kirwan, M.L., Daleo, P., Renzi, J.J., Butler, J., Osborne, T.Z., Nifong, J.C., and van de Koppel, J. 2019. Field experiments and meta-analysis reveal wetland vegetation as a crucial element in the coastal protection paradigm. Current Biology, 29, 1800-1806.

Renzi, J.J., He, Q., and Silliman, B.R. 2019. Harnessing positive species interactions to enhance coastal wetland restoration. Frontiers in Ecology and Evolution, 7: 131.

Silliman, B. R. & He, Q. 2018. Physical stress, consumer control, and new theory in ecology. Trends in Ecology & Evolution 33: 492-503.

Zhang, Y., Meng, H., Wang, Y. & He, Q. 2018. Herbivory enhances the resistance of mangrove forest to cordgrass invasion. Ecology 99: 1382-1390.

Thomsen, M. S., Altieri, A. H., Angelini, C., Bishop, M. J., Gribben, P. E., Lear, G., He, Q., Schiel, D. R., Silliman, B. R., South, P. M., Watson, D. M., Wernberg, T., & Zotz, G. 2018. Secondary foundation species enhance biodiversity. Nature Ecology & Evolution 2: 634–639.

Silliman, B. R., Hughes, B. B., Gaskins, L. C., He, Q., Tinker, M. T., Read, A., Nifong, J., & Stepp, R. 2018. Are the ghosts of nature’s past haunting ecology today? Current Biology 28: R532–R537.

Angelini, C., van Montfrans, S. G., Hensel, M. J. S., He, Q., & Silliman, B.R. 2018. The importance of an underestimated grazer under climate change: how crab density, consumer competition, and physical stress affect salt marsh resilience. Oecologia 187: 205–217.

Fariña, J. M., He, Q., Silliman, B. R., & Bertness, M.D. 2018. Biogeography of salt marsh plant zonation on the Pacific coast of South America. Journal of Biogeography 45: 238-247.

Silliman BR, BB Hughes, YS Zhang, & Q He. 2017. Business as usual leads to underperformance in coastal restoration. In: P Kareiva, M Marvier, & B Silliman (eds). Effective Conservation Science. Cambridge University Press, Cambridge, UK.

He, Q., Silliman, B. R., Liu, Z., & Cui, B. 2017. Natural enemies govern ecosystem resilience in the face of extreme droughts. Ecology Letters 20: 194–201.

          Research Highlight at Nature Climate Change

          Subject of an article at Yale Environment Review     

          Coverage: Science Daily, Science Newsline, Environmental News Network

He, Q., Silliman, B. R. & Cui, B. 2017. Incorporating thresholds into understanding salinity tolerance: A study using salt-tolerant plants in salt marshes. Ecology and Evolution 7: 6326-6333.

He, Q., & Silliman, B. R. 2016. Consumer control as a common driver of coastal vegetation worldwide. Ecological Monographs 86: 278–294.

         Featured on the cover


Silliman, B. R., Dixon, P. M., Wobus, C., He, Q., Daleo, P., Hughes, B. B., Willis, J. M., & Hester, M. W. 2016. Thresholds in marsh resilience to the Deepwater Horizon oil spill. Scientific Reports 6: 32520.


Liu, Z., Cui, B., & He, Q. 2016. Shifting paradigms in coastal restoration: Six decades' lessons from China. Science of the Total Environment 566: 205–214.


Freitas, R.F., Schrack, E.C., He, Q., Silliman, B.R., Furlong, E.B., Telles, A.C., & Costa, C.S.B. 2016. Consumer control of the establishment of marsh foundation plants in intertidal mudflats. Marine Ecology Progress Series 547: 79–89.


Fariña, J. M., He, Q., Silliman, B. R., & Bertness, M.D. 2016. Bottom-up and top-down human impacts interact to affect a protected coastal Chilean marsh. Ecology 97: 640–648.


Cui, B., He, Q., Gu, B, Bai, J., & Liu, X. 2016. China’s coastal wetlands: understanding environmental changes and human impacts for management and conservation. Wetlands 36: S1-S9.

       Introduction to the China Coastal Wetlands species issue


Silliman, B. R., Schrack, E. C., He, Q., Cope, R., Santoni, A., van der Heide, T., Jacobi, R., Jacobi, M., & van de Koppel, J. 2015. Facilitation shifts paradigms and can amplify coastal restoration efforts. Proceedings of the National Academy of Sciences USA 112: 14295–14300.

       Editor’s Choice at Science

       Coverage: Duke Today, NSF Discovery, Environmental Monitor


He, Q., & Silliman, B. R. 2015. Biogeographic consequences of nutrient enrichment for plant–herbivore interactions in coastal wetlands. Ecology Letters 18: 462–471.

       Coverage: Science Daily, Marine Pollution Bulletin, NSF Field News

       Cover Photo of the 2015 May issue of Ecology Letters


He, Q., Altieri, A.H., & Cui, B. 2015. Herbivory drives zonation of stress-tolerant marsh plants. Ecology 96: 1318–1328.


He, Q., & Cui, B. 2015. Multiple mechanisms sustain a plant-animal facilitation on a coastal ecotone. Scientific Reports 5: 8612.


He, Q., Bertness, M.D., Bruno, J.F., Li, B., Chen, G., Coverdale, T.C., Altieri, A.H., Bai, J., Sun, T., Pennings, S.C., Liu, J., Ehrlich, P.R., & Cui, B. 2014. Economic development and coastal ecosystem change in China. Scientific Reports 4: 5995.

       Editor’s Choice at Science

       Subject of an article at Scientific American

       Subject of an essay at The Conversation

       Coverage: Daily News, Science Newsline, Technobahn


He, Q., & Bertness, M.D.. 2014. Extreme stresses, niches, and positive species interactions along stress gradients. Ecology 95: 1437–1443.

       Featured on the cover

       ISI Highly Cited


He, Q., Bertness, M.D., & Altieri, A.H. 2013. Global shifts towards positive species interactions with increasing environmental stress. Ecology Letters 16: 695–706.

       Press release:

       Coverage:,, redOrbit

       Most cited Ecology Letters paper of 2013

       ISI Highly Cited


He, Q., Cui, B., Bertness, M.D., & An, Y. 2012. Testing the importance of plant strategies on facilitation using congeners in a coastal community. Ecology 93: 2023–2029.


He, Q., Cui, B.S., & An, Y. 2012. Physical stress, not biotic interactions, preclude an invasive grass from establishing in forb-dominated salt marshes. PLoS ONE 7: e33164.


He, Q., Chen, F.Y., Cui, B.S., & An, Y. 2012. Multi-scale segregations and edaphic determinants of marsh plant communities in a western Pacific estuary. Hydrobiologia 696: 171–183.


He, Q., Cui, B.S., & An, Y. 2011. The importance of facilitation in the zonation of shrubs along a coastal salinity gradient. Journal of Vegetation Science 22: 828–836.


Cui, B.S., He, Q., & An, Y. 2011. Spartina alterniflora invasions and effects on crab communities in a western Pacific estuary. Ecological Engineering 37: 1920–1924.


Cui, B.S., He, Q., Zhang, K.J., & Chen, X. 2011. Determinants of annual-perennial plant zonation across a salt-fresh marsh interface: a multistage assessment. Oecologia 166: 1067–1075.


Cui, B.S., He, Q., & An, Y. 2011. Community structure and abiotic determinants of salt marsh plant zonation vary across topographic gradients. Estuaries and Coasts 34: 459–469.


He, Q., Cui, B.S., Cai,Y.Z., Deng, J.F., Sun, T., & Yang, Z.F. 2009. What confines an annual plant to two separate zones along coastal topographic gradients? Hydrobiologia 630: 327–340.


Above is a partial list of my more recent publications.


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